Publications in TRACER 

  • Abbas, G., Jomaa, S., Bronstert, A., & Rode, M. (2023). Downstream changes in riverbank sediment sources and the effect of catchment size. Journal of Hydrology: Regional Studies, 46, 101340. https://doi.org/10.1016/j.ejrh.2023.101340
  • Abbas, G., Jomaa, S., Fink, P., Brosinsky, A., Nowak, K. M., Kümmel, S., Schkade, U., & Rode, M. (2024). Investigating sediment sources using compound-specific stable isotopes and conventional fingerprinting methods in an agricultural loess catchment. CATENA, 246, 108336. https://doi.org/10.1016/j.catena.2024.108336
  • Benisch, J., Helm, B., Chang, X., & Krebs, P. (2024). Can Short-Term Online-Monitoring Improve the Current WFD Water Quality Assessment Regime? Systematic Resampling of High-Resolution Data from Four Saxon Catchments. Water, 16(6), 889. https://doi.org/10.3390/w16060889
  • Benisch, J., Helm, B., Krauss, M., Byrne, H. A., Becker, S., Mayer, R. P., Rojas Gómez, K. L., Ahlheim, J., Brack, W., & Krebs, P. (2025). Fingerprints of micropollutants under baseflow and event discharge conditions: Analysing gradients along two urban streams. Water Science & Technology, 92(1), 34–52. https://doi.org/10.2166/wst.2025.091
  • Ben-Salem, N., Reinecke, R., Copty, N. K., Jaime Gómez-Hernández, J., Varouchakis, E. A., Karatzas, G. P., Rode, M., & Jomaa, S. (2023). Mapping steady-state groundwater levels in the Mediterranean region: The Iberian Peninsula as a benchmark. Journal of Hydrology, 626, 130207. https://doi.org/10.1016/j.jhydrol.2023.130207
  • Borriero, A., Kumar, R., Nguyen, T. V., Fleckenstein, J. H., & Lutz, S. R. (2023). Uncertainty in water transit time estimation with StorAge Selection functions and tracer data interpolation. Hydrology and Earth System Sciences, 27(15), 2989–3004. https://doi.org/10.5194/hess-27-2989-2023
  • Borriero, A., Musolff, A., Kumar, R., Fleckenstein, J. H., Lutz, S. R., & Nguyen, T. V. (2024). The value of instream stable water isotope and nitrate concentration data for calibrating a travel time‐based water quality model. Hydrological Processes, 38(5), e15154. https://doi.org/10.1002/hyp.15154
  • Borriero, A., Nguyen, T. V., Lutz, S. R., Fleckenstein, J. H., Musolff, A., & Kumar, R. (2024). Can the young water fraction reduce predictive uncertainty in water transit time estimations? Journal of Hydrology, 645, 132238. https://doi.org/10.1016/j.jhydrol.2024.132238
  • Chávez García Silva, R., Reinecke, R., Copty, N. K., Barry, D. A., Heggy, E., Labat, D., Roggero, P. P., Borchardt, D., Rode, M., Gómez-Hernández, J. J., & Jomaa, S. (2024). Multi-decadal groundwater observations reveal surprisingly stable levels in southwestern Europe. Communications Earth & Environment, 5(1), 387. https://doi.org/10.1038/s43247-024-01554-w
  • Chen, M., Jomaa, S., Lausch, A., Beudert, B., Ghaffar, S., Jia, W., & Rode, M. (2024). Impact of Forest Dieback on Hydrology and Nitrogen Export Using a New Dynamic Water Quality Model. Water Resources Research, 60(11), e2024WR037341. https://doi.org/10.1029/2024WR037341
  • Coder, L., Musolff, A., Kronsbein, P. M., Knöller, K., Büttner, O., Rinke, K., & Tittel, J. (2025). How anthropogenic modification of riverscapes reduces the resilience of floodplain water bodies to drought. Ecological Engineering, 219, 107686. https://doi.org/10.1016/j.ecoleng.2025.107686
  • Cromwell, L. A. (2023). Dark Waters: An Insight into Corporate Greed and PFAS Contamination. Ground Water, 61(3), 304–304. https://doi.org/10.1111/gwat.13302
  • Dehaspe, J., Sarrazin, F., Kumar, R., Fleckenstein, J. H., & Musolff, A. (2021). Bending of the concentration discharge relationship can inform about in-stream nitrate removal. Hydrology and Earth System Sciences, 25(12), 6437–6463. https://doi.org/10.5194/hess-25-6437-2021
  • Determann, M., Musolff, A., Frassl, M. A., Rinke, K., & Shatwell, T. (2023). Nutrient retention in a small reservoir under changed variability of inflow nutrient concentration. Inland Waters, 13(4), 560–575. https://doi.org/10.1080/20442041.2024.2305105
  • Ebeling, P. (2025). Groundwater droughts data for 6626 wells in Germany [Dataset]. HydroShare. https://doi.org/10.4211/hs.f0f661457b5e42288378b13ad0951d90
  • Ebeling, P., Dupas, R., Abbott, B., Kumar, R., Ehrhardt, S., Fleckenstein, J. H., & Musolff, A. (2021). Long-Term Nitrate Trajectories Vary by Season in Western European Catchments. Global Biogeochemical Cycles, 35(9), e2021GB007050. https://doi.org/10.1029/2021GB007050
  • Ebeling, P., Kumar, R., Weber, M., Knoll, L., Fleckenstein, J. H., & Musolff, A. (2021). Archetypes and Controls of Riverine Nutrient Export Across German Catchments. Water Resources Research, 57(4), e2020WR028134. https://doi.org/10.1029/2020WR028134
  • Ebeling, P., Musolff, A., Kumar, R., Hartmann, A., & Fleckenstein, J. H. (2024). Groundwater head responses to droughts across Germany. Copernicus GmbH. https://doi.org/10.5194/egusphere-2024-2761
  • Ehrhardt, S., Ebeling, P., Dupas, R., Kumar, R., Fleckenstein, J. H., & Musolff, A. (2021). Nitrate Transport and Retention in Western European Catchments Are Shaped by Hydroclimate and Subsurface Properties. Water Resources Research, 57(10), e2020WR029469. https://doi.org/10.1029/2020WR029469
  • Fárez‐Román, V., Rinke, K., Dunker, S., Hampel, H., & Shatwell, T. (2025). Phytoplankton community dynamics and vertical nutrient fluxes during the winter‐to‐spring transition in a monomictic temperate reservoir. Limnology and Oceanography, 70(6), 1678–1692. https://doi.org/10.1002/lno.70082
  • Große, A., Graeber, D., Fink, P., Reisinger, A. J., Kamjunke, N., Meyer, M., Ilić, M., Borchardt, D., & Perujo, N. (2025). Contrasting functional responses of benthic and hyporheic stream biofilms to light availability and macronutrient stoichiometry. Limnology and Oceanography, lno.70069. https://doi.org/10.1002/lno.70069
  •  Große, A., Perujo, N., Fink, P., Reisinger, A. J., Borchardt, D., McCarthy, M. J., & Graeber, D. (2025). Stream C to N to P ratios aligned with microbial needs enhance biofilm nitrate uptake and subsequent nitrogen loss. Biogeochemistry, 168(4), 62. https://doi.org/10.1007/s10533-025-01252-5
  • Große, A., Reisinger, A. J., Perujo, N., Fink, P., Borchardt, D., & Graeber, D. (2025). Acute changes in macronutrient stoichiometry alter nitrate uptake in benthic biofilms. Biogeochemistry: Rivers & Streams. https://doi.org/10.5194/egusphere-2025-3548
  • Harrow-Lyle, T. J., Lam, W. Y., Emilson, E. J. S., Mackereth, R. W., Mitchell, C. P. J., & Melles, S. J. (2023). Watershed characteristics and chemical properties govern methyl mercury concentrations within headwater streams of boreal forests in Ontario, Canada. Journal of Environmental Management, 345, 118526. https://doi.org/10.1016/j.jenvman.2023.118526
  • Heinemann, N., Yang, S., Büttner, O., & Borchardt, D. (2025). Nutrient loading and stream order shape benthic and pelagic spring algal biomass in a large, temperate river basin (Elbe River). Journal of Environmental Management, 383, 125440. https://doi.org/10.1016/j.jenvman.2025.125440
  • Hildebrandt, S., Krueger, E. H., Ruhl, A. S., & Borchardt, D. (2024). Efficacy of point source legislation quantified for a 64-year river water quality trajectory of phosphorus loading. Journal of Environmental Management, 352, 119956. https://doi.org/10.1016/j.jenvman.2023.119956
  • Holden, M. M., Timshina, A., Mehdi, Q., Cromwell, L. A., Osborne, T., Aufmuth, J., & Bowden, J. A. (2024). Have per- and polyfluoroalkyl substances (PFAS) infiltrated Florida’s freshwater springs? Science of The Total Environment, 952, 175826. https://doi.org/10.1016/j.scitotenv.2024.175826
  • Hubig, A., Musolff, A., Shatwell, T., Weitere, M., Wachholz, A., Ebeling, P., & Scharfenberger, U. (2025). Linking spatial patterns of chlorophyll a and phosphorus concentrations: River length and upstream lakes control realized eutrophication in German rivers. Water Research, 287, 124372. https://doi.org/10.1016/j.watres.2025.124372
  • Jähkel, A., Graeber, D., Fleckenstein, J. H., & Schmidt, C. (2022). Hydrologic Turnover Matters—Gross Gains and Losses of Six First‐Order Streams Across Contrasting Landscapes and Flow Regimes. Water Resources Research, 58(7), e2022WR032129. https://doi.org/10.1029/2022WR032129
  •  Javed, A., Arnillas, C. A., Saha, R., Chang, R., & Arhonditsis, G. B. (2025). Striking a balance between food security and water quality: A modelling analysis of agricultural management practices in mitigating non-point source phosphorus pollution. Science of The Total Environment, 974, 179135. https://doi.org/10.1016/j.scitotenv.2025.179135
  • Javed, A., Cheng ,Vincent Y. S., & and Arhonditsis, G. B. (2019). Detection of spatial and temporal hydro-meteorological trends in Lake Michigan, Lake Huron and Georgian Bay. Aquatic Ecosystem Health & Management, 22(1), 1–14. https://doi.org/10.1080/14634988.2018.1500850
  • Javed, A., Loudoun, C., Neumann, A., Arnillas, C. A., Okoli, A., & Arhonditsis, G. B. (2024a). Connecting past, present, and future trends of hydraulic and phosphorus loading in the Bay of Quinte tributaries, Ontario, Canada. Journal of Hydrology: Regional Studies, 53, 101818. https://doi.org/10.1016/j.ejrh.2024.101818
  • Javed, A., Loudoun, C., Neumann, A., Arnillas, C. A., Okoli, A., & Arhonditsis, G. B. (2024b). Connecting past, present, and future trends of hydraulic and phosphorus loading in the Bay of Quinte tributaries, Ontario, Canada. Journal of Hydrology: Regional Studies, 53, 101818. https://doi.org/10.1016/j.ejrh.2024.101818
  • Javed, A., Neumann, A., Cai, H., Alberto Arnillas, C., & Arhonditsis, G. B. (2024). A reservoir-based approach of the SWAT hydrological model in the Napanee River and Wilton Creek agricultural watersheds, Bay of Quinte. Journal of Great Lakes Research, 50(5), 102404. https://doi.org/10.1016/j.jglr.2024.102404
  • Javed, A., Saber, A., Neumann, A., Arnillas, C. A., & Arhonditsis, G. B. (2025). Characterization of biogeochemical cycles in agricultural watersheds: Integrating regional modelling assessment with downstream water quality. Journal of Environmental Management, 393, 127086. https://doi.org/10.1016/j.jenvman.2025.127086
  • Kong, X., Ghaffar, S., Determann, M., Friese, K., Jomaa, S., Mi, C., Shatwell, T., Rinke, K., & Rode, M. (2022). Reservoir water quality deterioration due to deforestation emphasizes the indirect effects of global change. Water Research, 221, 118721. https://doi.org/10.1016/j.watres.2022.118721
  • Lam, W. Y., Mackereth, R. W., & Mitchell, C. P. J. (2022). Landscape controls on total mercury and methylmercury export from small boreal forest catchments. Biogeochemistry, 160(1), 89–104. https://doi.org/10.1007/s10533-022-00941-9
  • Lam, W. Y., Mackereth, R. W., & Mitchell, C. P. J. (2024a). Landscape characteristics govern the impacts of beaver ponds on surface water methylmercury concentrations in boreal watersheds. Environmental Research: Ecology, 3(2), 025004. https://doi.org/10.1088/2752-664X/ad4f93
  • Lam, W. Y., Mackereth, R. W., & Mitchell, C. P. J. (2024b). Mercury concentrations and export from small central Canadian boreal forest catchments before, during, and after forest harvest. Science of The Total Environment, 912, 168691. https://doi.org/10.1016/j.scitotenv.2023.168691
  • Li, Y., Yang, X., Lischeid, G., Wollheim, W. M., Jomaa, S., Zhou, X., & Rode, M. (2025). Responses of Wetted River Network     Contraction and Expansion Dynamics to Prolonged Drought. Water Resources Research, 61(6), e2024WR038938. https://doi.org/10.1029/2024WR038938
  • Liu, J., Wang, S., Jia, W., Chen, M., Wang, X., Yao, H., & Ni, X. (2023). Impact of Climate Change on Reservoir Operation during the Dry Season in the Pearl River Basin. Water, 15(21), Article 21. https://doi.org/10.3390/w15213749
  • Mangal, V., Lam, W. Y., Huang, H., Emilson, E. J. S., Mackereth, R. W., & Mitchell, C. P. J. (2022). Molecular correlations of dissolved organic matter with inorganic mercury and methylmercury in Canadian boreal streams. Biogeochemistry, 160(1), 127–144. https://doi.org/10.1007/s10533-022-00944-6
  • Meyer, M., Koschorreck, M., Weitere, M., Kneis, D., & Perujo, N. (2024). Dissolved organic matter quality, hydrological connectivity and microbial activity shape phosphorus buffering in river-floodplain systems. Science of The Total Environment, 957, 177452. https://doi.org/10.1016/j.scitotenv.2024.177452
  • Nakulopa, F., Bärlund, I., & Borchardt, D. (2024). How a reservoir modulates downstream water quality under declining upstream loading and progressing climate change. Science of The Total Environment, 912, 169460. https://doi.org/10.1016/j.scitotenv.2023.169460
  • Nicholls, C., & Hoffmann, T. (2025). Cross-Sectional Variability of Suspended Sediment Concentration in the Rhine River. Water, 17(23), 3391. https://doi.org/10.3390/w17233391
  • Pasqualini, J., Graeber, D., Bartusch, A., Kümmel, S., Hernandez, Z. L. D., Musat, N., Sunjidmaa, N., Weitere, M., & Brauns, M. (2024). Disentangling effects of multiple agricultural stressors on benthic and hyporheic nitrate uptake. Biogeochemistry, 167(3), 287–299. https://doi.org/10.1007/s10533-024-01130-6
  • Pasqualini, J., Majdi, N., Weitere, M., & Brauns, M. (2025). The contribution of the hyporheos to whole-stream invertebrate secondary production. Freshwater Science, 44(2), 170–186. https://doi.org/10.1086/735823
  • Peter, L. G., & Lee, L. S. (2025). Sources and Pathways of PFAS Occurrence in Water Sources: Relative Contribution of La    nd-Applied Biosolids in an Agricultural Dominated Watershed. Environmental Science & Technology, 59(2), 1344–1353. https://doi.org/10.1021/acs.est.4c09490
  • Peter, L., Modiri-Gharehveran, M., Alvarez-Campos, O., Evanylo, G. K., & Lee, L. S. (2025). PFAS fate using lysimeters during degraded soil reclamation using biosolids. Journal of Environmental Quality, 54(1), 41–53. https://doi.org/10.1002/jeq2.20576
  • Rojas-Gómez, K. L., Binder, M., Walther, M., & Engelmann, C. (2023). A parsimonious approach to predict regions affected by sewer-borne contaminants in urban aquifers. Environmental Monitoring and Assessment, 195(12), 1517. https://doi.org/10.1007/s10661-023-12027-6
  • Rouhani, A., Ben-Salem, N., D’Oria, M., Silva, R. C. G., Viglione, A., Copty, N. K., Rode, M., Barry, D. A., Gómez-Hernández, J. J., & Jomaa, S. (2025). Direct impact of climate change on groundwater levels in the Iberian Peninsula. Science of The Total Environment, 970, 179009. https://doi.org/10.1016/j.scitotenv.2025.179009
  • Saavedra, F. A., Musolff, A., Von Freyberg, J., Merz, R., Basso, S., & Tarasova, L. (2022). Disentangling scatter in long-term concentration–discharge relationships: The role of event types. Hydrology and Earth System Sciences, 26(23), 6227–6245. https://doi.org/10.5194/hess-26-6227-2022
  • Saavedra, F., Musolff, A., Von Freyberg, J., Merz, R., Knöller, K., Müller, C., Brunner, M., & Tarasova, L. (2024). Winter post-droughts amplify extreme nitrate concentrations in German rivers. Environmental Research Letters, 19(2), 024007. https://doi.org/10.1088/1748-9326/ad19ed
  • Saavedra, F., Vergopolan, N., Musolff, A., Merz, R., Wang, Z., Winter, C., & Tarasova, L. (2025). From Soil Moisture Spatial Patterns to Catchment Nitrate Dynamics using Explainable AI. Preprints. https://doi.org/10.22541/essoar.175855522.20150676/v1
  • Schauer, L. S., Jawitz, J. W., Cohen, M. J., & Musolff, A. (2025). Spatial and Temporal Variability of River Water Quality. Hydrological Processes, 39(5), e70154. https://doi.org/10.1002/hyp.70154
  • Shin, Y., Reisinger, A. J., Flint, M., Salinas, T., Martin, J. B., & Cohen, M. J. (2025). Nutrient Limitation Induces a Productivity Decline From Light‐Controlled Maximum. Journal of Geophysical Research: Biogeosciences, 130(4), e2024JG008597. https://doi.org/10.1029/2024JG008597
  • Sunjidmaa, N., Mendoza-Lera, C., Hille, S., Schmidt, C., Borchardt, D., & Graeber, D. (2022). Carbon limitation may override fine-sediment induced alterations of hyporheic nitrogen and phosphorus dynamics. Science of the Total Environment, 837, 155689. https://doi.org/10.1016/j.scitotenv.2022.155689
  • Sunjidmaa, N., Mendoza-Lera, C., Pasqualini, J., Fink, P., Bartusch, A., Borchardt, D., Jähkel, A., & Graeber, D. (2025). Irradiance and biofilm age control daytime and nighttime macronutrient cycling in stream mesocosms. Biogeochemistry, 168(2), 25. https://doi.org/10.1007/s10533-025-01215-w
  • Teran-Velasquez, G., Helm, B., & Krebs, P. (2022). Longitudinal River Monitoring and Modelling Substantiate the Impact of Weirs on Nitrogen Dynamics. Water, 14(2), Article 2. https://doi.org/10.3390/w14020189
  • Teran-Velasquez, G., Helm, B., & Krebs, P. (2023). High Spatiotemporal Model-Based Tracking and Environmental Risk-Exposure of Wastewater-Derived Pharmaceuticals across River Networks in Saxony, Germany. Water, 15(11), Article 11. https://doi.org/10.3390/w15112001
  • Wachholz, A., Dehaspe, J., Ebeling, P., Kumar, R., Musolff, A., Saavedra, F., Winter, C., Yang, S., & Graeber, D. (2023). Stoichiometry on the edge—Humans induce strong imbalances of reactive C:N:P ratios in streams. Environmental Research Letters, 18(4), 044016. https://doi.org/10.1088/1748-9326/acc3b1
  • Wachholz, A., Jawitz, J. W., & Borchardt, D. (2024). From Iron Curtain to green belt: Shift from heterotrophic to autotrophic nitrogen retention in the Elbe River over 35 years of passive restoration. Biogeosciences, 21(15), 3537–3550. https://doi.org/10.5194/bg-21-3537-2024
  • Wachholz, A., Jawitz, J. W., Büttner, O., Jomaa, S., Merz, R., Yang, S., & Borchardt, D. (2022). Drivers of multi-decadal nitrate regime shifts in a large European catchment. Environmental Research Letters, 17(6), 064039. https://doi.org/10.1088/1748-9326/ac6f6a
  • Wang, H.-J., Merz, R., & Basso, S. (2025). Constructing a geography of heavy-tailed flood distributions: Insights from common streamflow dynamics. Hydrology and Earth System Sciences, 29(6), 1525–1548. https://doi.org/10.5194/hess-29-1525-2025
  • Wang, H.-J., Merz, R., Yang, S., & Basso, S. (2023). Inferring heavy tails of flood distributions through hydrograph recession analysis. Hydrology and Earth System Sciences, 27(24), 4369–4384. https://doi.org/10.5194/hess-27-4369-2023
  • Wang, H.-J., Merz, R., Yang, S., Tarasova, L., & Basso, S. (2023). Emergence of heavy tails in streamflow distributions: The role of spatial rainfall variability. Advances in Water Resources, 171, 104359. https://doi.org/10.1016/j.advwatres.2022.104359
  • Winter, C., Jawitz, J. W., Ebeling, P., Cohen, M. J., & Musolff, A. (2024). Divergence Between Long-Term and Event-Scale Nitrate Export Patterns. Geophysical Research Letters, 51(10), e2024GL108437. https://doi.org/10.1029/2024GL108437
  • Winter, C., Lutz, S. R., Musolff, A., Kumar, R., Weber, M., & Fleckenstein, J. H. (2021). Disentangling the Impact of Catchment Heterogeneity on Nitrate Export Dynamics From Event to Long‐Term Time Scales. Water Resources Research, 57(1), e2020WR027992. https://doi.org/10.1029/2020WR027992
  • Zhang, C., Huang, Y., Javed, A., & Arhonditsis, G. B. (2019). An ensemble modeling framework to study the effects of climate change on the trophic state of shallow reservoirs. Science of The Total Environment, 697, 134078. https://doi.org/10.1016/j.scitotenv.2019.134078
  • Zhang, X., Yang, X., Jomaa, S., & Rode, M. (2020). Analyzing impacts of seasonality and landscape gradient on event-scale nitrate-discharge dynamics based on nested high-frequency monitoring. Journal of Hydrology, 591, 125585. https://doi.org/10.1016/j.jhydrol.2020.125585
  • Zill, J., Perujo, N., Fink, P., Mallast, U., Siebert, C., & Weitere, M. (2024). Contribution of groundwater-borne nutrients to eutrophication potential and the share of benthic algae in a large lowland river. Science of The Total Environment, 951, 175617. https://doi.org/10.1016/j.scitotenv.2024.175617
  • Zill, J., Siebert, C., Rödiger, T., Schmidt, A., Gilfedder, B. S., Frei, S., Schubert, M., Weitere, M., & Mallast, U. (2023). A way to determine groundwater contributions to large river systems: The Elbe River during drought conditions. Journal of Hydrology: Regional Studies, 50, 101595. https://doi.org/10.1016/j.ejrh.2023.101595
  • Zill, J., Suckow, A., Mallast, U., Sültenfuß, J., Schmidt, A., & Siebert, C. (2025). Will groundwater-borne nutrients affect river eutrophication in the future? A multi-tracer study along the Elbe River. Hydrology and Earth System Sciences, 29(23), 6885–6900. https://doi.org/10.5194/hess-29-6885-2025