title

Research for the Environment

German

Dr. Ingolf Kühn

Kontakt/Adresse

Dr. Ingolf Kühn
Senior Scientist

Department Community Ecology
Helmholtz Centre for
Environmental Research - UFZ
Theodor-Lieser-Str. 4
06120 Halle, Germany

Tel: (+49) 345 558 5311
Fax: (+49) 341 235 45 5311
ingolf.kuehn@ufz.de

Research interests

I am vegetation scientist with some experience in floristics, taxonomy and systematics in my past life. Now I went for something completely different: The primary work at the department deals with plant invasions and their relevance for the diversity of plant species. Therefore, I analyse land use patterns, environmental parameters and plant species distribtion on a macroecological scale. Another focus are biological and ecological traits of invasive plants. Thus I model and administrate the "Database on Biological and Ecological Traits of the Flora of Germany (BiolFlor) that was prepared by many collaborators at our department. Additionally, I am interested in patterns and processes of plant diversity in urbanised areas.
Recently, I broadened up my area of research to include distribution patterns under global change.
Methodologically, I am interested in methods to account for spatial and phylogenetic non-independence (autocorrelation).

During my academic career at the Ruhr-University of Bochum at the chair of systematic botany I worked mainly in database related projects, e. g. mapping schemes, checklist on the flora of Germany, and the Information System of Botanical Gardens in Germany (SysTax). However, my research focussed on vegetation science. My PhD thesis was about a comparison of ancient and recent forests in Westphalia in the importance of forest continuity vs soil characteristcs. As I am also highly interested in birds, I wrote my diploma thesis about the conservation value of a former military training area, regarding birds, flora, and vegetation.

Current Projects

Regional scenarios, combined effects and management options (Cluster 1 in Topic 1 "Land Use Options - Strategies and Adaptation to Global Change" of the project oriented funding of the Helmholtz Association), cluster speaker.
Regionalised impacts of climate change on experimental species models and species ranges for Germany (Project 1 in Cluster 2 "Impacts of Climate Change on Ecological Patterns and Processes" of Topic 1 (see above)), project leader.
BiolFlor: a database on biological and ecological traits of the flora of Germany
ALARM: Assessing LArge-scale environmental Risks with tested Methods (Integrated project funded by the European Union under FP 6), Deputy Coordinator
Protected areas in Germany under climate change - risks and management options (Schutzgebiete Deutschlands im Klimawandel – Risiken und Handlungsoptionen) (Funded by the Federal Agency for Nature Conservation), collaborator
COCONUT: understanding effeCts Of land use Changes ON ecosystems to halt loss of biodiversity due to habitat destrUction, fragmenTation and degradation. (Scientific support to policies project funded by the European Union under FP 6), collaborator
Climate and Fauna: Impacts of recent climate change on the fauna in Germany (Auswirkungen des rezenten Klimawandels auf die Fauna in Deutschland)(Funded by the Federal Agency for Nature Conservation), collaborator.

Past Projects

MACIS: Minimisation of and Adaptation to Climate change Impacts on biodiverSity (Scientific support to policies project funded by the European Union under FP 6), Coordinator
Flora & Climate: Modelling the effect of climate change on the flora of Germany (Modellierung der Auswirkungen des Klimawandels auf die Flora) (Funded by the Federal Agency for Nature Conservation), Coordinator
Virtual Institute Macroecology (Funded by the Helmholtz Association of National Research Centres, Scientific Secretary
DAISIE: Delivering Alien Invasive Species Inventories for Europe (Specific targeted research project funded by the European Union under FP 6), collaborator
INVASIONS: Das Invasionspotential gebietsfremder Arten - Identifikation, Bewertung, Risikomanagement (The Invasion Potential of Alien Species - Identification, Assessment, and Risk Management) (Funded by the Federal Ministry for Education and Research under the BioTEAM programme), Deputy Coordinator
LEDA: A database on the life history of the Northwest European flora (Funded by the European Union under FP 5)
Analysis of distribution patterns of species and functional groups in the UFZ research cluster "Biological invasions, dispersal processes and distribution patterns", project coordination
Urban biodiversity in the former UFZ research cluster "Urban ecology and urban development", collaborator

Selected publications

Complete list of publications

Settele, J., Kudrna, O., Harpke, A., Kühn, I., van Swaay, C., Verovnik, R., Warren, M., Wiemers, M., Hanspach, J., Hickler, T., Kühn, E., van Halder, I., Veling, K., Vliegenhart, A., Wynhoff, I., Schweiger, O. 2008. Climatic Risk Atlas of European Butterflies. BioRisk 1. Pensoft, Sofia (Bulgaria).
Online version
Dormann, C.F., Kühn, I. 2008. Statistische Analyse biologischer Daten (mit dem freien Programmpaket R). Skript, 2. Auflage. Download

Accepted, in press

Bräuniger, C., Knapp, S., Kühn, I., Klotz, S. Testing taxonomic and landscape surrogates for biodiversity in an urban setting. Landscape and Urban Planning, accepted.
Chytrý, M., Wild, J., Pyšek, P., Jarošík, V., Dendoncker, N., Reginster, I., Pino, J.,Maskell, L.C., Vilà, M., Kühn, I., Spangenberg, J.H., Settele, J. Projecting trends in plant invasions in Europe under different scenarios of future land-use change. Global Ecology & Biogeography, accepted.
Hickler, T., Vohland, K, Miller, P.A:, Smith, B., Costa, L., Feehan, J., Giesecke, T., Fronzek, S.,Carter, T., Cramer, W., Kühn, I., Sykes, M.T. Climate-driven changes in European potential natural vegetation in relation to the Natura 2000 protected area network. Global Ecology and Biogeography, accepted.
Pompe, S., Hanspach, J., Badeck, F.-W., Klotz, S., Bruehlheide, H., Kühn, I.. Investigating habitat-specific co-occurrence of plant species under climate change. Basic and Applied Ecology, accepted.
Schweiger, O., Biesmeijer, J.C., Bommarco, R., Hickler, T., Hulme, P. E., Klotz, S., Kühn, I., Moora, M., Nielsen, A., Ohlemüller, R., Petanidou, T., Potts, S. G., Pyšek, P., Stout, J., Sykes, M. T., Tscheulin, T., Vilà, M., Walther, G.-R., Westphal, C, Winter, M., Zobel, M., Settele, J. Multiple stressors on biotic interactions: how climate change and alien species interact to affect pollination. Biological Reviews, Early View (doi: 10.1111/j.1469-185X.2010.00125.x). Reprint
Schweiger, O., Heikkinen, R.K., Harpke, A., Hickler, T., Klotz, S., Kudrna, O., Kühn, I., Pöyry, J., Settele, J. Increasing range mismatching of interacting species under global change is related to species traits. Global Ecology and Biogeography, accepted.
Spangenberg, J., Carter, T., Fronzek, S., Jaeger, J., Jylhä, K., Kühn, I., Omann, I., Paul, A., Reginster, I., Rounsevell, M., Schweiger, O., Stocker, A., Sykes, M., Settele, J. Scenarios for investigating risks to biodiversity. Global Ecology and Biogeography, accepted.

2010

Bergmann, J., Pompe, S., Ohlemüller, R., Freiberg, M., Klotz, S., Kühn, I. 2010. The Iberian Peninsula as a potential source for the plant species pool in Germany under projected climate change. Plant Ecology 207: 191-201. Reprint , Supplement
Bierman, S. M., Butler, A., Marion, G., Kühn, I. 2010. Bayesian image restoration models for combining expert knowledge on recording activity with species distribution data. Ecography 33: 451-460. Reprint
Carl, G., Kühn, I. 2010. A wavelet-based extension of generalized linear models to remove the effect of spatial autocorrelation. Geographical Analysis 42: 323-337. Reprint
Fuentes, N., Ugarte, E., Kühn, I., Klotz, S. 2010. Alien plants in Southern South America. A framework for evaluation and management of mutual risk of invasion between Chile and Argentina. Biological Invasions 12: 3227–3236. Reprint
Hammen, V.C., Biesmeijer, J.C., Bommarco, R., Budrys, E., Christensen, T.R., Fronzek, S., Grabaum, R., Jaksic, P., Klotz, S., Kramarz, P., Kröel-Dulay, G., Kühn, I., Mirtl, M., Moora, M., Petanidou, T., Pino, J., Potts, S.G., Rortais, A., Schulze, C.H., Steffan-Dewenter, I., Stout, J., Szentgyörgyi, H., Vighi, M., Vujic, A., Westphal, C., Wolf, T., Zavala, G., Zobel, M., Settele, J., Kunin, W.E. 2010. Establishment of a cross-European field site network in the ALARM project for assessing large-scale changes in biodiversity. Environmental Monitoring & Assessment 164: 337-348. Reprint
Hanspach, I., Kühn, I., Pompe, S., Klotz, S. Predictive performance of plant species distribution models depends on species traits. Perspectives in Plant Ecology, Evolution and Systematics 12: 219–225. Reprint
Knapp, S., Kühn, I., Stolle, J. & Klotz, S. Changes in the functional composition of a Central European urban flora over three centuries. Perspectives in Plant Ecology, Evolution and Systematics 12: 235-244. Reprint
Küster, E.C., Durka, W., Kühn, I., Klotz, S. 2010. Differences in trait compositions of non-indigenous and native plants across Germany. Biological Invasions 12: 2001-2012. Reprint
Pyšek, P., Jarošík, V., Hulme,P.E., Kühn, I., Wild, J., Arianoutsou, M., Bacher, S., Chiron, F., Didžiulis, V., Essl, F., Genovesi, P., Gherardi, F., Hejda, M., Kark, S., Lambdon, P.W., Desprez-Loustau, M.-L., Nentwig, W., Pergl, J., Poboljšaj, K., Rabitsch, W., Roques, A., Roy, D.B., Shirley, S., Solarz, W., Vilá, M., Winter, M. 2010. Disentangling the role of environmental and human pressures on biological invasions across Europe. Proceedings of the National Academy of Sciences 107: 12157-12162. Reprint
Vilà, M., Basnou, C., Pyšek, P., Josefsson, M., Genovesi, P., Gollasch, S., Nentwig, W., Olenin, S., Roques, A., Roy, D., Hulme, P.E., DAISIE partners (Adriopoulos, P., Arianoutsou, M., Augustin, S., Baccetti, N., Bacher, S., Bacon, J., Bazos, I., Bolshagin, P., Bretagnolle, F., Chiron, F., Clergeau, P., Cochard, P.O., Cocquempot, C., Coeur d’Acier, A., Cooper, J., Daunys, D., David, M., Delipetrou, P., Didžiulis, V., Dorkeld, F., Essl, F., Galil, B., Gasquez, J., Georghiou, K., Gudžinskas, Z., Hatzofe, O., Hejda, M., Hill, M., Jarošík, V., Kark, S., Klotz, S., Kobelt, M., Kokkoris, Y., Kotarac, M., Kühn, I., Lambdon, P., Lange, E., Lopez-Vaamonde, C., Loustau, M.-L., Marcer, A., Martinez, M., McLoughlin, M., Migeon, A., Minchin, D., Navajas, M., Navajas, P., Olenina, I., Ostler, R., Ovcharenko, I., Panov, V.E., Papacharalambous, E., Pascal, M., Pergl, J., Perglová, I., Phillipov, A., Pino, J., Poboljsaj, K., Rabitsch, W., Rasplus, J.-Y., Rodionova, N., Roy, H., Sauvard, D., Scalera, R., Schwartz, A., Sedláček, O., Shirley, S., Trocchi, V., Winter, M., Yart, A., Yiannitsaros, A., Zagatti, P., Zikos, A.). 2010. How well do we understand the impacts of alien species on ecosystem services? A pan-European cross-taxa assessment. Frontiers in Ecology and the Environment 8:135-144. Reprint
Winter, M., Kühn, I., La Sorte, F., Schweiger, O., Nentwig, W., Klotz, S. 2010 The role of non-native plants and vertebrates in defining patterns of compositional dissimilarity within and across continents. Global Ecology and Biogeography 19: 332–342. Reprint , Supporting Information

2009

Knapp, S., Kühn, I., Bakker, J.P., Kleyer, M., Klotz, S., Ozinga, W.A., Poschlod, P., Thompson, K., Thuiller, W., Römermann, C. 2009. How species traits and affinity to urban land use control large-scale species frequency. Diversity & Distributions 15: 533-546. Reprint
Kühn, I., Nobis, M.P., Durka, W. 2009. Combining spatial and phylogenetic filtering in a trait analysis of plant flowering phenology in Switzerland. Global Ecology & Biogeography 18: 745-758. Reprint , Supplement
Pyšek, P, Jarošík, V., Pergl, J., Randall, R., Chytrý, M., Kühn, I.,Tichý, L., Danihelka, J., Chrtek jun., J., Sádlo, J. 2009. The global invasion success of Central European plants is related to distribution characteristics in their native range and species traits. Diversity & Distributions 15: 891-903. Reprint
Pyšek, P., Lambdon, P.W., Arianoutsou, M., Kühn, I., Pino, J., Winter, M. 2009. Aliens vascular plants of Europe. In DAISIE (eds.), The handbook of alien species in Europe, 43-61. Springer, Dordrecht. Reprint
Seppelt, R., Kühn, I., Klotz, S., Kabisch, S., Schloter, M., Auge, H., Görg, C., Frank, K., Jax, K. 2009. Land Use Options - Strategies and Adaptation to Global Change. GAIA - Ecological Perspectives for Science and Society 18/1: 77-80. Reprint
Walther, G.-R., Roques, A., Hulme, P. E., Sykes, M. T., Pyšek, P., Kühn, I., Zobel, M., Bacher, S., Botta-Dukát, Z., Bugmann, H., Czúcz, B., Dauber, J., Hickler, T., Jarošík, V., Kenis, M., Klotz, S., Minchin, D., Moora, M., Nentwig, W., Ott, J., Panov, V. E., Reineking, B., Robinet, C., Semenchenko, V., Solarz, W., Thuiller, W., Vilà, M., Vohland, K., Settele, J. 2009. Alien species in a warmer world - risks and opportunities. Trends in Ecology & Evolution 24: 686-693. Reprint
Williamson, M., Dehnen-Schmutz, K., Kühn, I., Hill, M., Klotz, S., Milbau, A., Stout, J., Pyšek, P. 2009. The distribution of range sizes of native and alien plants in four European countries and the effects of residence time. Diversity & Distributions 15: 158-166. Reprint
Winter, M.,Schweiger, O., Klotz, S., Nentwig, W., Andriopoulos, P., Arianoutsou, M., Basnou, C., Delipetrou, P., Didziulis, V., Hejda, M., Hulme, P.E., Lambdon, P., Pergl, P., Pyšek, P., Roy, D., Kühn, I. 2009. Plant extinctions and introductions lead to phylogenetic and taxonomic homogenization of the European flora. Proceedings of the National Academy of Sciences 106: 21721-21725. Open Access.

2008

Brändle, M., Kühn, I., Klotz, S., Belle, C., Brandl, R. 2008. Species richness of herbivores on exotic host plants increases with time since introduction of the host. Diversity & Distributions 14: 905-912. Reprint , Supplements
Carl, G., Dormann, C., Kühn, I. 2008. A wavelet-based method to remove spatial autocorrelation in the analysis of species distributional data. Web Ecology 8: 22-29 Reprint , R Codes
Carl, G., Kühn, I. 2008. Analyzing spatial ecological data using linear regression and wavelet analysis. Stochastic Environmental Research and Risk Assessment 22: 315-324. Reprint
Fuentes, N., Kühn, I., Ugarte, E., Klotz, S. 2008. Alien plants in Chile. Inferring invasion periods from herbarium records. Biological Invasions 10: 649-657. Reprint
Götzenberger, L., Kühn, I., Klotz, S. 2008. Effects of habitat disturbance and pollination type on the interspecific variation in pollen-ovule ratios. Preslia 80: 423-437. Reprint , Appendix
Hanspach, J., Kühn, I., Pyšek, P., Boos, E., Klotz, S. 2008. Correlates of naturalization and occupancy of introduced ornamentals in Germany. Perspectives in Plant Ecology, Evolution and Systematics 10: 241-250. Reprint
Hulme, P.E., Bacher, S., Kenis, M., Klotz, S., Kühn, I., Minchin, D., Nentwig, W., Olenin, O., Panov, V., Pergl, J., Pyšek, P., Roques, A., Sol, D., Solarz, W. & Vilá, M. 2008: Grasping at the routes of biological invasions: a framework for integrating pathways into policy. Journal of Applied Ecology, 45: 403–414. Reprint
Kleyer, M., Bekker, R.M., Knevel, I.C., Bakker, J.P., Thompson, K., Sonnenschein, M., Poschlod, P., van Groenendael, J.M., Klimeš, L., Klimešová, J., Klotz, S., Rusch, G.M., Hermy, M., Adriaens, D., Boedeltje, G., Bossuyt, B., Dannemann, A., Endels, P., Götzenberger, L., Hodgson, J.G., Jackel, A-K., Kühn, I., Kunzmann, D.,Ozinga, W.A., Römermann, C., Stadler, M., Schlegelmilch, J., Steendam, H.J., Tackenberg, O., Wilmann, B., Cornelissen, J.H.C., Eriksson, O., Garnier, E., & Peco, B. 2008. The LEDA Traitbase: A database of plant life-history traits of North West European flora. Journal of Ecology 96:1266-1274. Reprint , Supplement
Knapp, S., Kühn, I., Mosbrugger, V., Klotz, S. 2008. Do protected areas in urban and rural landscapes differ in species diversity? Biodiversity & Conservation 17:1595-1612. Reprint
Knapp, S., Kühn, I., Schweiger, O., Klotz, S. 2008. Challenging urban species diversity: contrasting phylogenetic patterns across plant functional groups in Germany. Ecology Letters 11: 1054-1064. Reprint
Knapp, S., Kühn, I., Wittig, R., Ozinga, W., Poschlod, P., Klotz, S. 2008. Urbanisation causes shifts of species’ trait state frequencies. Preslia 80: 375-388. Reprint
Kühn, I., Böhning-Gaese, K., Cramer, W., Klotz, S. 2008. Macroecology meets global change research. Global Ecology & Biogeography 17: 3-4. Reprint
Kühn, I., Sykes, M.T., Berry, P.M., Thuiller, W., Piper, J.M., Nigmann, U., Araújo, M.B., Balletto, E., Bonelli, S., Cabeza, M., Guisan, A., Hickler, T., Klotz, S., Metzger, M., Midgley, G., Musche, M., Olofsson, J., Paterson, J.S., Penev, L., Rickebusch, S., Rounsevell, M.D.A.R., Schweiger, O., Wilson, E., Settele, J. 2008. MACIS: Minimisation of and Adaptation to Climate change Impacts on biodiverSity. GAIA - Ecological Perspectives for Science and Society 17/4: 393-395. Reprint
Küster, E. C., Kühn, I., Bruelheide, H., Klotz, S. 2008. Trait interactions help explain plant invasion success in the German flora. Journal of Ecology 96: 860-868. Reprint , Supplement
featured as Editor's choice in Journal of Ecology 96(5) Editor's choice
Lambdon, P, Pyšek, P., Basnou, C., Arianoutsou, M., Ess, F., Hejda, M., Jarošík, V., Pergl, J., Winter, M., Anastasiu, P., Andriopoulos, P., Bazos, I., Brundu, G., Celesti-Grapow, L., Chassot, P., Delipetrou, P., Josefsson, M., Kark, S., Klotz, S., Kokkoris, Y., Kühn, I., Marchante, H., Perglová, I., Pino, J., Vilà, M., Zikos, A., Roy, D., Hulme, P. 2008. Alien flora of Europe: species diversity, temporal trends, geographical patterns and research needs. Preslia 80: 101-149. Reprint , Supplement
Liira, L., Schmidt, T., De Cock, R., Tsipe A., Arens, P., Augenstein, I., Bailey, D., Billeter, R., Bukáček, R., Burel, F., De Blust, G., De Cock, R., Dirksen, J., Edwards, P. J., Hamerský, R., Herzog, F., Klotz, S., Kühn, I., Le Coeur, D., Miklová, P., Roubalova, M., Schweiger, O., Smulders, M.J.M., van Wingerden, W.K.R.E., Bugters, R., Zobel, M. 2008: Plant functional group composition and large-scale species richness in the agricultural landscapes of Europe. Journal of Vegetation Science 19: 3-14. Reprint
Lososová, Z., Chytrý, M., Kühn, I. 2008. Plant attributes determining regional commonness and rarity of weeds on Central European arable land. Journal of Biogeography 35: 177-187. Reprint , Supplement
Pompe, S., Badeck, F.-W., Hanspach, J., Klotz, S., Thuiller, W., Kühn, I. 2008. Projecting impact on plant distributions under climate change - a case study from Germany. Biology Letters 4: 564-567. Reprint , Supplement
Pyšek, P., Lambdon, P.W., Arianoutsou, M., Kühn, I., Pino, J., Winter, M. 2009. Aliens vascular plants of Europe. In DAISIE (eds.), The handbook of alien species in Europe, 43-61. Springer, Dordrecht. Reprint
Qian, H., McKinney, M.L., Kühn, I. 2008. Effects of introduced species on floristic similarity: comparing two US states. Basic and Applied Ecology 9: 617-625. Reprint , Supplement
Schweiger, O., Klotz, S., Durka, W., Kühn, I. 2008. A comparative test of phylogenetic diversity indices. Oecologia 157:485-495. Reprint
Schweiger, O., Settele, J., Kudrna, O., Klotz, S., Kühn, I. 2008. Climate change can cause spatial mismatch of trophically interacting species. Ecology 89: 3472–3479. Reprint , Appendices
Settele, J., Spangenberg, J, Kühn, I. 2008. Large projects can create useful partnerships. Nature 453: 850. Reprint
Sol, D., Vilá, M., Kühn, I. 2008. The comparative analysis of historical alien introductions. Biological Invasions 10: 1119-1129. Reprint
Tautenhahn, S., Heilmeier,H., Götzenberger, L., Klotz, S., Wirth, C., Kühn, I. 2008. On the biogeography of seed mass in Germany - distribution patterns and environmental correlates. Ecography 31: 457-468. Reprint , Appendix
Winter, M., Kühn, I. , Nentwig, W., Klotz, S. 2008. Spatial aspects of trait homogenization within the German Flora. Journal of Biogeography 12: 2289 - 2297. Reprint , Supplement

2007

Carl, G., Kühn, I. 2007. Analyzing spatial autocorrelation in species distributions using Gaussian and Logit models. Ecological Modelling, 207(2-4): 159-170. Reprint , R-code
Dormann, C. F., McPherson, J. M., Araújo, M. B., Bivand, R., Bolliger, J., Carl, G., Davies, R. G., Hirzel, A., Jetz, W., Kissling, W. D., Kühn, I., Ohlemüller, R., Peres-Neto, P. R., Reineking, B., Schröder, B., Schurr, F. M., Wilson, R. 2007. Methods to account for spatial autocorrelation in the analysis of species distributional data: a review. Ecography 30: 609-628. Reprint , R-codes , data
Götzenbeger, L., Durka, W., Kühn, I., Klotz, S. 2007. The relationship of pollen-ovule ratio and seed size – another comparative test of a sex allocation hypothesis. Evolutionary Ecology Research 9: 1145-1161. Reprint
Kühn, I. 2007. Incorporating spatial autocorrelation may invert observed patterns. Diversity & Distributions, 13: 66-69. Reprint , electronic appendix
Kühn, I. and Klotz, S. 2007. From Ecosystem invasibility to local, regional and global patterns of invasive species. In Nentwig, W. (ed.), Biological invasions. Ecological Studies 193: 181-196. Berlin, Heidelberg, New York: Springer. Reprint
Settele, J, Kühn, I., Klotz, S., Hammen, V., Spangenberg, J. 2007. Is the EC afraid of its own visions? Science 315: 1220. Reprint

2006

Götzenberger, L., Durka, W., Kühn, I., Klotz, S. 2006. The relationship of pollen-ovule ratio and seed size – a comparative test of a sex allocation hypothesis. Evolutionary Ecology Research, 8: 1101-1116. Reprint
Kühn, I., Bierman, S. M., Durka, W. and Klotz, S. 2006. Relating geographical variation in pollination types to environmental and spatial factors using novel statistical methods. New Phytologist, 172: 127-139. Reprint , electronic appendix
Kühn, I. and Klotz, S. 2006. Urbanisation and homogenization - comparing the floras of urban and rural areas in Germany. Biological Conservation, 127: 292-300. Reprint
Lososová, Z., Chytrý, M., Kühn, I., Hájek, O., Horáková, V., Pyšek, P., Tichý, L. 2006. Patterns of plant traits in annual vegetation of man-made habitats in Central Europe. Perspectives in Plant Ecology, Evolution and Systematics, 8: 69-81. Reprint
Wania, A., Kühn, I., Klotz, S. 2006. Biodiversity patterns of plants in agricultural and urban landscapes in Central Germany - spatial gradients of species richness. Landscape and Urban Planning, 75(1): 97-110. Reprint

pre 2006

Settele, J., Hammen, V., Hulme, P.E., Karlson, U., Klotz, S., Kotarac, M., Kunin, W.E., Marion, G., O'Connor, M., Petanidou, T., Petersen, K., Potts, S., Pritchard, H., Pyšek, P., Rounsevell, M., Spangenberg, J., Steffan-Dewenter, I., Sykes, M.T., Vighi, M., Zobel, M. and Kühn, I. 2005. ALARM: Assessing LArge scale environmental Risks for biodiversity with tested Methods. GAIA - Ecological Perspectives for Science and Society, 14(1): 69-72. Reprint

Kühn, I., Brandenburg, M. and Klotz, S. 2004. Why do alien plant species that reproduce in natural habitats occur more frequently? Diversity & Distributions, 10(5/6): 417-425. Reprint
Kühn, I., Durka, W. and Klotz, S. 2004. BiolFlor – a new plant-trait database as a tool for plant invasion ecology. Diversity & Distributions, 10(5/6): 363-365. Reprint , electronic appendix
Kühn, I., Brandl, R. and Klotz, S. 2004. The flora of German cities is naturally species rich. Evolutionary Ecology Research, 6: 749-764. Reprint
Deutschewitz, K., Lausch, A., Kühn, I. and Klotz, S. 2003. Native and alien plant species richness in relation to spatial heterogeneity on a regional scale in Germany. Global Ecology and Biogeography, 12: 299-311. Reprint

Kühn, I., May, R., Brandl, R. and Klotz, S. 2003. Plant distribution patterns in Germany - Will aliens match natives? Feddes Repertorium, 114: 559-573. Reprint
Kühn, I. and Klotz, S. 2003. The alien flora of Germany - basics from a new German database. In Plant invasions: ecological threats and management solutions. (L.E. Child, J.H. Brock, G. Brundu, K. Prach, P. Pyšek, P.M. Wade and M. Williamson, eds.), pp. 89-100. Leiden: Backhuys. Reprint

Klotz, S., Kühn, I. and Durka, W. 2002 (eds.). BIOLFLOR – Eine Datenbank zu biologisch-ökologischen Merkmalen der Gefäßpflanzen in Deutschland. Schriftenreihe für Vegetationskunde 38. Bonn: Bundesamt für Naturschutz. 334 S.