species information (Systematics, BiolFlor-Names, Status) |
BiolFlor-Name |
family |
german name(s) |
Echium vulgare L. |
Boraginaceae » |
Gewöhnlicher Natternkopf
|
reference |
Kühn, I., Klotz, S. 2002: Systematik, Taxonomie und Nomenklatur. In: Klotz, S., Kühn, I., Durka, W. (Hrsg.): BIOLFLOR - Eine Datenbank mit biologisch-ökologischen Merkmalen zur Flora von Deutschland. Schriftenreihe für Vegetationskunde. Bonn »
|
|
Synonymy (taxonomy after Wisskirchen & Haeupler, 1998) |
Echium vulgare L. |
accepted |
|
Chromosome number » |
Basic chromosome number |
8 |
Chromosome number 2n |
32 |
Chromosome number (Germany) |
True |
Most common chromosome race |
False |
Ploidy level |
4 (tetraploid) |
Ploidy type |
P (polyploid with haploid basic number) |
Reference |
Oberdorfer, E. 1983: Pflanzensoziologische Exkursionsflora. Ulmer, Stuttgart |
|
Life form » |
Life form |
H (Hemicryptophyte) |
Qualifier |
always [1] |
Life span » |
Life span |
b (biennial) |
Qualifier |
always [1] |
Life span |
h (pluriennial-hapaxanthic) |
Qualifier |
sometimes [2] |
Storage organs » |
Storage organs |
r (Primary storage root) |
Qualifier |
sometimes [2] |
Root metamorphoses » |
Root metamorphoses |
r (Primary storage root) |
Qualifier |
sometimes [2] |
Rosettes » |
Rosettes |
h (Hemirosette plant) |
Qualifier |
always [1] |
special references:
|
Meusel, H. 1952: Über Wuchsform, Verbreitung und Phylogenie einiger mediterran mitteleuropäischer Angiospermen-Gattungen. Flora 139:333-393
|
Hensen, I. 1997: Life strategies of xerothermic grasslands - mechanisms of reproduction and colonization within Stipetum capillatae s.l. and Adonido-Brachypodietum pinnati. Feddes Repertorium 108:425-452
|
Frey, W., Hensen, I. 1995: Spergulo morisonii-Corynephoretum canescentis (Frühlingsspark-Silbergrasflur) - Lebensstrategien von Binnendünen- und Lockersandbesiedlern. Feddes Repertorium 106:533-553
|
Klemow, K.M., Raynal, D.J. 1985: Demography of two facultative biennial plant species in an unproductive habitat. Journal of Ecology 73:147-167
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van Breemen, A.M.M. 1984: Comparative germination ecology of three short lived monocarpic Boraginaceae. Acta Botanica Neerlandica 33:283-305
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Ebel, F., Mühlberg, H. 1987: Vergleichend-ökomorphologische Untersuchungen an Pflanzen-Taxa eines Trocken- und Feuchtbiotops. Hercynia N. F. 24:408-424
|
Hensen, I., Kentrup, M. 1998: Teucrio botryos-Melicetum ciliatae (Traubegamander-Wimpergrasflur) - Lebensstrategien in einer xerothemen Fels-Pioniergesellschaft. Tuexenia 18:217-236
|
Kästner, A., Karrer, G. 1995: Übersicht der Wuchsformentypen als Grundlage für deren Erfassung in der "Flora von Österreich". Florae Austriacae Novit. 3:1-51
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Salisbury, E. 1952: Downs and dunes. Their plant life and its environment. Bells & Sons, London
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Flowering phenology » |
Begin of flowering (months) |
5 |
End of flowering (months) |
7 |
Duration of flowering (months) |
3 |
Number of flowering phases |
0 |
Symphenological Groups » |
Phenological group |
6 (Cornus sanguinea-Melica uniflora-Phase) Phenological season: start of early summer |
Reference |
Dierschke, H 1995: Phänologische und symphänologische Artengruppen von Blütenpflanzen in Mitteleuropa. Tuexenia 15: 523-560 |
|
Type of reproduction » |
Type of reproduction |
s (by seed/by spore) |
Reference |
Krumbiegel, A. 2002: Morphologie der vegetativen Organe (außer Blätter). In: Klotz, S., Kühn, I., Durka, W. (Hrsg.) BiolFlor - Biologisch-ökologische Daten zur Flora Deutschlands, S. 1-2, Bonn |
Type of seed production » |
Type of seed production |
s (amphimictic, sexual) |
Reference |
Knuth, P. 1898: Handbuch der Blütenbiologie. Leipzig |
Dicliny » |
Dicliny |
so (hermaphroditic) |
Abundance |
the rule [2] |
Reference |
Knuth, P. 1898: Handbuch der Blütenbiologie. Leipzig |
Dicliny |
gd (gynodioecious) |
Abundance |
rare [4] |
Reference |
Knuth, P. 1898: Handbuch der Blütenbiologie. Leipzig |
Dichogamy » |
Dichogamy |
a2 (protandrous) |
Reference |
Knuth, P. 1898: Handbuch der Blütenbiologie. Leipzig |
Self-sterility and self-incompatibility » |
Self-sterility and self-incompatibility |
SC (self-compatibel) |
Reference |
Mulligan, G.A. 1972: Autogamy, allogamy, and pollination in some Canadian weeds. Canadian Journal of Botany 50:1767-1771 |
Systematic reference |
species [S] |
Pollen vector » |
Pollen vector |
in (insects) Pollen vector group: zo (Tierbestäubung) |
Abundance |
the rule [2] |
Reference |
Knuth, P. 1898: Handbuch der Blütenbiologie. Leipzig |
Systematic reference |
species [S] |
Breeding system » |
Breeding system |
XF (facultative allogamous) |
Reference |
Rademaker, M.C.J., de Jong, T.J., van der Meijden, E. 1999: Selfing rates in natural populations of Echium vulgare: a combined empirical and model approach. Functional Ecology 13:828-837 |
P/O ratio » |
P/O ratio |
20313.0 |
Reference |
Rademaker, M.C.J., Dejong, T.J., Klinkhamer, P.G.L. 1997: Pollen dynamics of bumble-bee visitation on Echium vulgare. Functional Ecology 11:554-563 |
Floral rewards » |
Floral rewards |
1 (nectar) |
Reference |
Knuth, P. 1898: Handbuch der Blütenbiologie. Leipzig |
Amount of floral reward |
4 (plenty) |
Flower color » |
Flower colour |
at first pink, then bright blue |
Basic flower colour |
ro (pink) |
Reference |
Schubert, R., Jäger, E.J., Werner, K. (Hrsg.) 1987: Exkursionsflora Band 3. Atlas der Gefäßpflanzen. 6. Aufl., Volk und Wissen, Berlin |
UV reflection of the flower » |
UV reflection in the periphery of the flower |
5 (40-66 %) |
UV reflection in the centre of the flower |
5 (40-66 %) |
UV reflection pattern |
0 (no UV reflectance patterns) |
Flower class after MUELLER » |
Flower class after MUELLER |
H (hymenoptere flowers) Typical pollinators: hymenopteres |
Flower type after KUGLER » |
Flower type after KUGLER |
5.2 (lip flowers, throat flowers) |
|
Fruit type » |
Fruit type |
BrF (lomentum) |
Diaspore type » |
Diaspore type |
FZb (fruit with appendage) |
Diaspore type |
TF (mericarp) |
Germinule type » |
Heteromorphic type |
n (not heteromorphic) |
Germinule type |
FZb (fruit with appendage) |
Information available |
True |
Development of nutritious appendages |
|
Development of flat appendages |
|
Development of elongated appendages |
always [1] |
Development of additional appendages |
always [1] |
Heteromorphic type |
n (not heteromorphic) |
Germinule type |
TF (mericarp) |
Information available |
True |
Development of nutritious appendages |
|
Development of flat appendages |
|
Development of elongated appendages |
|
Development of additional appendages |
|
Weights » |
Measured object |
Diaspore |
Type of germinule or diaspore |
TF (mericarp) |
Constraint |
False |
Heteromorphic type |
n (not heteromorphic) |
Weight (minimum) [mg] |
2.2 |
Weight (mean) [mg] |
2.9 |
Weight (maximum) [mg] |
3.4 |
Measured object |
Germinule |
Type of germinule or diaspore |
TF (mericarp) |
Constraint |
False |
Heteromorphic type |
n (not heteromorphic) |
Weight (minimum) [mg] |
2.2 |
Weight (mean) [mg] |
2.9 |
Weight (maximum) [mg] |
3.4 |
Sizes » |
Measured object |
Germinule |
Type of germinule or diaspore |
TF (mericarp) |
Constraint |
False |
Heteromorphic type |
n (not heteromorphic) |
Length (minimum) [mm] |
2.1 |
Length (mean) [mm] |
2.7 |
Length (maximum) [mm] |
3.0 |
Width (minimum) [mm] |
1.2 |
Width (mean) [mm] |
1.8 |
Width (maximum) [mm] |
2.2 |
Height (minimum) [mm] |
1.0 |
Height (mean) [mm] |
1.6 |
Height (maximum) [mm] |
2.0 |
Measured object |
Diaspore |
Type of germinule or diaspore |
TF (mericarp) |
Constraint |
False |
Heteromorphic type |
n (not heteromorphic) |
Length (minimum) [mm] |
2.1 |
Length (mean) [mm] |
2.7 |
Length (maximum) [mm] |
3.0 |
Width (minimum) [mm] |
1.2 |
Width (mean) [mm] |
1.8 |
Width (maximum) [mm] |
2.2 |
Height (minimum) [mm] |
1.0 |
Height (mean) [mm] |
1.6 |
Height (maximum) [mm] |
2.0 |
special references:
|
Grime, J.P., Mason, G., Curtis, A.V., Rodman, J., Band, S.R., Mowforth, M.A., Neal, A.M., Shaw, S. 1981: A comparative study of germination characteristics in a local flora. Journal of Ecology 69:1017-1069 (Gattung)
|
Briemle, G. 1996: Farbatlas Kräuter und Gräser. Ulmer, Stuttgart (Gattung)
|
Beijerink, W. 1947: Zadenatlas der Nederlandsche Flora. Biol. Stat. Wijster, Wageningen (Gattung)
|
Böttner, I., Frey, W., Hensen, I. 1997: Carex humilis-Gesellschaft im unteren Unstruttal (mitteldeutsches Trockengebiet) - Lebensstrategien einer xerothermen Vegetationseinheit. Feddes Repertorium 108:7; 583-8; 602 (Gattung)
|
Brouwer, W., Stählin, A. 1955: Handbuch der Samenkunde. DLG-Verlag, Frankfurt/Main (Gattung)
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Hanf, M. 1984: Ackerunkräuter Europas mit ihren Keimlingen und Samen. . BLV Verlagsgesellschaft, München, Wien, Zürich. (Gattung)
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Luftensteiner, H.W. 1982: Untersuchungen zur Verbreitungssbiologie von Pflanzengemeinschaften an vier Standorten in Niederösterreich. Bibliotheca Botanica 135:1-68 (Gattung)
|
Heinisch, O. 1955: Samenatlas der wichtigsten Futterpflanzen und ihrer Unkräuter. Deutsche Akad. Landwirtsch.-Wiss., Berlin (Gattung)
|
Hensen, I., Kentrup, M. 1998: Teucrio botryos-Melicetum ciliatae (Traubegamander-Wimpergrasflur) - Lebensstrategien in einer xerothemen Fels-Pioniergesellschaft. Tuexenia 18:217-236 (Gattung)
|
|
Grassland utilization indicator values » |
Occurrence |
N (fallow lands) |
Guild |
K (forb) |
Mowing tolerance |
2 (between 1 and 3 (intolerant to sensitive to mowing)) |
Grazing tolerance |
3 (sensitive to grazing) |
Trampling tolerance |
2 (between 1 and 3 (intolerant to sensitive to trampling)) |
Foraging value |
2 (no or little forage value) |
Foraging value for deer |
|
|
Habitat » |
Habitat |
T10.3.1 (null) Alliance: ???
|
Occurrence questionable |
False |
Occurence restricted |
False |
Annotation |
|
Habitat |
T10.3.2 (null) Alliance: ???
|
Occurrence questionable |
False |
Occurence restricted |
False |
Annotation |
|
Habitat |
T10.4.5 (null) Alliance: ???
|
Occurrence questionable |
False |
Occurence restricted |
False |
Annotation |
|
Habitat |
T6.2 (null) Alliance: Festuco-Brometea
|
Occurrence questionable |
False |
Occurence restricted |
False |
Annotation |
lückig |
Habitat |
T6.2.1.4 (null) Alliance: ???
|
Occurrence questionable |
False |
Occurence restricted |
False |
Annotation |
|
Habitat |
T8.2.2.2 (null) Alliance: Echio-Melilotion
|
Occurrence questionable |
False |
Occurence restricted |
False |
Annotation |
|
|
Class » |
English circumscription |
04.4 (Communities of heavy metal soils) Class: Violetea calaminariae
|
English circumscription |
05.2 (Pioneer communities of hair-grasses) Class: Koelerio-Corynephoretea
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English circumscription |
05.3 (Fescue-brome communities of dry and semi-dry grasslands) Class: Festuco-Brometea
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English circumscription |
06.4 (Annual ruderal communities) Class: Sisymbrietea officinalis
|
English circumscription |
06.5 (Eusiberian ruderal mugwort and thistles communities, and couch-grasslands) Class: Artemisietea vulgaris
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